Cheese-and-Hamburger Casserole

Ingredients

• 1 pound ground round
• 1 cup chopped onion
• 3 garlic cloves, crushed
• 1 (8-ounce) package presliced mushrooms
• 6 tablespoons tomato paste
• 1 teaspoon sugar
• 1 teaspoon dried thyme
• 1 teaspoon dried oregano
• 1/4 teaspoon pepper
• 1 (28-ounce) can whole tomatoes, undrained and chopped
• 1/3 cup all-purpose flour
• 2 1/2 cups 2% low-fat milk
• 1 cup (4 ounces) crumbled feta cheese
• 3/4 cup (3 ounces) shredded part-skim mozzarella cheese
• 4 cups uncooked penne (tubular-shaped pasta)
• 1 tablespoon chopped fresh parsley (optional)