Dungeness Crab Mac 'n' Cheese with Sourdough Breadcrumbs - Dough Wines

Ingredients

• 3/4 cup fresh sourdough breadcrumbs
• 1/4 cup grated Parmesan cheese
• 1 tablespoon olive oil
• 1/2 teaspoon kosher salt
• A generous pinch of freshly ground black pepper
• 1 pound medium pasta shells or tubes (such as conchiglie, penne, or cavatappi)
• 1/4 cup ½ stick unsalted butter, plus more for greasing
• 1/4 cup all-purpose flour
• 4 cups whole milk (warmed)
• 1 1/2 cups sharp white Cheddar cheese (grated )
• 1 1/2 cups Gruyère cheese (grated )
• 1/2 cup Parmesan cheese (grated )
• 1 teaspoon kosher salt (plus more to taste)
• 1/4 teaspoon freshly ground black pepper (plus more to taste)
• 1 tablespoon fresh lemon zest
• A pinch cayenne
• 1 pound cooked Dungeness crab meat (picked over to remove any bits of shell)
• Chopped chives (for garnish)
• Lemon wedges (for serving)